We Here See At The Bottom Of The Cliffs, Beds Containing Sharks' Teeth And Sea-Shells Of Extinct Species, Passing Above Into An Indurated Marl, And From That Into The Red Clayey Earth Of The Pampas, With Its Calcareous Concretions And The Bones Of Terrestrial Quadrupeds.

This vertical section clearly tells us of a large bay of pure salt- water, gradually encroached on, and at last converted into the bed of a muddy estuary, into which floating carcasses were swept.

At Punta Gorda, in Banda Oriental, I found an alternation of the Pampaean estuary deposit, with a limestone containing some of the same extinct sea-shells; and this shows either a change in the former currents, or more probably an oscillation of level in the bottom of the ancient estuary. Until lately, my reasons for considering the Pampaean formation to be an estuary deposit were, its general appearance, its position at the mouth of the existing great river the Plata, and the presence of so many bones of terrestrial quadrupeds: but now Professor Ehrenberg has had the kindness to examine for me a little of the red earth, taken from low down in the deposit, close to the skeletons of the mastodon, and he finds in it many infusoria, partly salt-water and partly fresh-water forms, with the latter rather preponderating; and therefore, as he remarks, the water must have been brackish. M. A. d'Orbigny found on the banks of the Parana, at the height of a hundred feet, great beds of an estuary shell, now living a hundred miles lower down nearer the sea; and I found similar shells at a less height on the banks of the Uruguay; this shows that just before the Pampas was slowly elevated into dry land, the water covering it was brackish. Below Buenos Ayres there are upraised beds of sea-shells of existing species, which also proves that the period of elevation of the Pampas was within the recent period.

In the Pampaean deposit at the Bajada I found the osseous armour of a gigantic armadillo-like animal, the inside of which, when the earth was removed, was like a great cauldron; I found also teeth of the Toxodon and Mastodon, and one tooth of a Horse, in the same stained and decayed state. This latter tooth greatly interested me, [3] and I took scrupulous care in ascertaining that it had been embedded contemporaneously with the other remains; for I was not then aware that amongst the fossils from Bahia Blanca there was a horse's tooth hidden in the matrix: nor was it then known with certainty that the remains of horses are common in North America. Mr. Lyell has lately brought from the United States a tooth of a horse; and it is an interesting fact, that Professor Owen could find in no species, either fossil or recent, a slight but peculiar curvature characterizing it, until he thought of comparing it with my specimen found here: he has named this American horse Equus curvidens. Certainly it is a marvellous fact in the history of the Mammalia, that in South America a native horse should have lived and disappeared, to be succeeded in after- ages by the countless herds descended from the few introduced with the Spanish colonists!

The existence in South America of a fossil horse, of the mastodon, possibly of an elephant, [4] and of a hollow-horned ruminant, discovered by MM. Lund and Clausen in the caves of Brazil, are highly interesting facts with respect to the geographical distribution of animals. At the present time, if we divide America, not by the Isthmus of Panama, but by the southern part of Mexico [5] in lat. 20 degs., where the great table-land presents an obstacle to the migration of species, by affecting the climate, and by forming, with the exception of some valleys and of a fringe of low land on the coast, a broad barrier; we shall then have the two zoological provinces of North and South America strongly contrasted with each other. Some few species alone have passed the barrier, and may be considered as wanderers from the south, such as the puma, opossum, kinkajou, and peccari. South America is characterized by possessing many peculiar gnawers, a family of monkeys, the llama, peccari, tapir, opossums, and, especially, several genera of Edentata, the order which includes the sloths, ant-eaters, and armadilloes. North America, on the other hand, is characterized (putting on one side a few wandering species) by numerous peculiar gnawers, and by four genera (the ox, sheep, goat, and antelope) of hollow-horned ruminants, of which great division South America is not known to possess a single species. Formerly, but within the period when most of the now existing shells were living, North America possessed, besides hollow-horned ruminants, the elephant, mastodon, horse, and three genera of Edentata, namely, the Megatherium, Megalonyx, and Mylodon. Within nearly this same period (as proved by the shells at Bahia Blanca) South America possessed, as we have just seen, a mastodon, horse, hollow- horned ruminant, and the same three genera (as well as several others) of the Edentata. Hence it is evident that North and South America, in having within a late geological period these several genera in common, were much more closely related in the character of their terrestrial inhabitants than they now are. The more I reflect on this case, the more interesting it appears: I know of no other instance where we can almost mark the period and manner of the splitting up of one great region into two well- characterized zoological provinces. The geologist, who is fully impressed with the vast oscillations of level which have affected the earth's crust within late periods, will not fear to speculate on the recent elevation of the Mexican platform, or, more probably, on the recent submergence of land in the West Indian Archipelago, as the cause of the present zoological separation of North and South America. The South American character of the West Indian mammals [6] seems to indicate that this archipelago was formerly united to the southern continent, and that it has subsequently been an area of subsidence.

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